National "111" Center for Cellular Regulation and Molecular Pharmaceutics, Key Laboratory of Fermentation Engineering (Ministry of Education), Hubei Key Laboratoy of Industrial Microbiology, Hubei Research Center of Food Fermentation Engineering and Technology, Hubei University of Technology, Wuhan 430068, Hubei, China. Electronic address: wuqian@hbut.edu.cn. National "111" Center for Cellular Regulation and Molecular Pharmaceutics, Key Laboratory of Fermentation Engineering (Ministry of Education), Hubei Key Laboratoy of Industrial Microbiology, Hubei Research Center of Food Fermentation Engineering and Technology, Hubei University of Technology, Wuhan 430068, Hubei, China. State Key Laboratory of Marine Resource Utilization in South China Sea/ Ministry of Education, Key Laboratory of Food Nutrition and Functional Food of Hainan Province/ Engineering Research Center of Utilization of Tropical Polysaccharide Resources/ School of Food Science and Engineering, Hainan University, Haikou, China. Electronic address: guozq@hainanu.edu.cn. State Key Laboratory of Marine Resource Utilization in South China Sea/ Ministry of Education, Key Laboratory of Food Nutrition and Functional Food of Hainan Province/ Engineering Research Center of Utilization of Tropical Polysaccharide Resources/ School of Food Science and Engineering, Hainan University, Haikou, China. National "111" Center for Cellular Regulation and Molecular Pharmaceutics, Key Laboratory of Fermentation Engineering (Ministry of Education), Hubei Key Laboratoy of Industrial Microbiology, Hubei Research Center of Food Fermentation Engineering and Technology, Hubei University of Technology, Wuhan 430068, Hubei, China. Electronic address: njfeng@hbut.edu.cn.