College of Life Sciences, Northwest A&F University, Yangling, Shaanxi, 712100, China. Electronic address: lijsh@nwafu.edu.cn. College of Life Sciences, Northwest A&F University, Yangling, Shaanxi, 712100, China. School of Environmental Science and Engineering, Shaanxi University of Science & Technology, Xi'an, Shaanxi, 710021, China. School of Environmental Science and Engineering, Shaanxi University of Science & Technology, Xi'an, Shaanxi, 710021, China. Electronic address: jiahonglei@sust.edu.cn.