National Key Laboratory of Crop Genetic Improvement and National Centre of Plant Gene Research, Huazhong Agricultural University, Wuhan 430070, China. Key Laboratory of Arable Land Conservation (Middle and Lower Reaches of Yangtze River), Ministry of Agriculture, Huazhong Agricultural University, Wuhan 430070, China. Plant and Crop Sciences Division, School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, United Kingdom. Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing 100081, China. School of Agriculture, Policy and Development, University of Reading, Reading RG6 6AR, United Kingdom. Southern Cross Plant Science, Southern Cross University, Lismore NSW 2480, Australia. The James Hutton Institute, Invergowrie, Dundee DD2 5DA, United Kingdom. King Saud University, Riyadh 11451, Kingdom of Saudi Arabia.