Key Laboratory of Vegetation Ecology, Institute of Grassland Science, Northeast Normal University, Changchun 130024, Jilin Province, China; Key Laboratory of Mollisols Agroecology, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, Jilin Province, China. Key Laboratory of Mollisols Agroecology, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, Jilin Province, China. Key Laboratory of Vegetation Ecology, Institute of Grassland Science, Northeast Normal University, Changchun 130024, Jilin Province, China. Electronic address: gaoyz108@nenu.edu.cn. Institute of Research and Development, Duy Tan University, 03 Quang Trung, Da Nang, Viet Nam; Signaling Pathway Research Unit, RIKEN Center for Sustainable Resource Science, Yokohama, Kanagawa, Japan. Electronic address: son.tran@riken.jp. Key Laboratory of Mollisols Agroecology, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, Jilin Province, China. Electronic address: tiancj@neigae.ac.cn.