Robert W Holley Center for Agriculture and Health, USDA-ARS, Cornell University, Ithaca, New York, 14853, USA; Plant Breeding and Genetics Section, School of Integrative Plant Science, Cornell University, Ithaca, New York, 14853, USA. Robert W Holley Center for Agriculture and Health, USDA-ARS, Cornell University, Ithaca, New York, 14853, USA; Plant Breeding and Genetics Section, School of Integrative Plant Science, Cornell University, Ithaca, New York, 14853, USA. Electronic address: ll37@cornell.edu.