Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universität München, 80333 Munich, Germany. GeoBio-Center, Ludwig-Maximilians-Universität München, 80333 Munich, Germany. Department of Biology, University of Konstanz, 78457 Constance, Germany. Department of Biology, Philipps-Universität, 35032 Marburg, Germany. Institute for Molecular Evolution, Heinrich Heine Universität Düsseldorf, 40225 Düsseldorf, Germany.